Summer schools, courses, online resources and textbooks across structural biology techniques.
Courses, free online material and books, from first steps to specialist methods. Use the buttons to show one technique.
CCP4 and the British Crystallographic Association
Every two years in September (most recently 6–11 September 2026, John Innes Centre, Norwich; next in 2028) · UK
in person · introductory
For students and researchers relatively new to crystallography. Lectures cover protein expression through to structure validation, with supervised computer tutorials and a crystal manipulation and cryo-cooling workshop.
CCP4
Every January · UK
in person · all
Annual meeting on computational methods in macromolecular structure determination, with a different theme each year.
CCP-EM
Courses through the year
hybrid · all
Courses and workshops on cryo-EM data processing and model building from the UK's collaborative computational project for cryo-EM.
EMBO
Many courses each year, across Europe
in person · intermediate
Hands-on courses, including regular ones in structural biology methods; search the listings by topic.
EMBL-EBI
Live courses and on-demand material
hybrid · all
Training on bioinformatics and structural data resources such as PDBe, EMDB and AlphaFold DB.
CCPBioSim
Workshops through the year
hybrid · all
Training in biomolecular simulation from the UK collaborative computational project.
Instruct-ERIC
Courses through the year
in person · intermediate
Training courses across structural biology techniques at Instruct centres in Europe.
Free courses, tutorials and resources.
Caltech · Free video lecture course.
RCSB PDB · Educational resources on structural biology and the PDB.
EMBL-EBI
BioXTAS RAW
CSIC
Cambridge Institute for Medical Research
Bernhard Rupp
University of the Basque Country
University of York
EPFL
Wolfram
Outline of Crystallography for Biologists
David Blow · Oxford University Press, 2002
Crystallography Made Crystal Clear: A Guide for Users of Macromolecular Models
Gale Rhodes · Academic Press, 1999
Three-Dimensional Electron Microscopy of Macromolecular Assemblies
Joachim Frank · Oxford University Press, 2nd edition, 2006
Small Angle X-Ray and Neutron Scattering from Solutions of Biological Macromolecules
Dmitri Svergun, Michel Koch, Peter Timmins and Roland May · Oxford University Press, 2013
Fundamentals of Crystallography
C. Giacovazzo, H.L. Monaco, G. Artioli, D. Viterbo, M. Milanesio, G. Gilli, G. Zanotti, G. Ferraris and M. Catti · IUCr Texts on Crystallography, Oxford University Press, 2011
Protein Crystallography
T.L. Blundell and L.N. Johnson · Academic Press, 1976
Macromolecular Crystallography with Synchrotron Radiation
J.R. Helliwell · Cambridge University Press, 1992
Principles of Protein X-ray Crystallography
J. Drenth · Springer, 2007
X-Ray Crystallography of Biomacromolecules: A Practical Guide
Albrecht Messerschmidt · Wiley, 2007
Macromolecular Crystallography: Conventional and High-Throughput Methods
M.R. Sanderson and J.V. Skelly · Oxford University Press, 2007
B. Rupp · Garland Science, 2009
Crystals, X-rays and Proteins: Comprehensive Protein Crystallography
D. Sherwood and J. Cooper · Oxford University Press, 2010 and 2015 (erratum and extra material at crxp.org.uk)
Protein Crystallography: Methods and Protocols
A. Wlodawer, Z. Dauter and M. Jaskolski · Springer, 2017